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- ArticleGraf JD, Karch F, Moreillon MC.Experientia. 1977 Dec 15;33(12):1582-4.Investigations of the green frogs from western Europe for electrophoretic variations at 4 enzyme loci demonstrated a new form which must be considered as a hybrid between Rana ridibunda and R. perezi. Biochemical evidence supports the hypothesis that its reproduction is hybridogenetic, as it is for R. esculenta.